Firefighters and emergency responders operate in dynamic environments where uncertain responsibilities and acute stress can compromise coordinated action. This technical report develops an integrated, testable framework linking psychological safety, role ambiguity and adaptive team performance under high-stress incident conditions. An analytically grounded review synthesizes research on team psychological safety, role stress, adaptation, incident command, situational awareness and occupational demands. A prospective multiteam, multiwave design is specified to distinguish individual perceptions from team-level functioning and to reduce common-method bias. The proposed model assesses direct effects, the psychological-safety × role-ambiguity interaction, incident-stress contingency, and indirect pathways through communication and coordination. Multilevel structural equation modelling, measurement invariance, reliability tests, prespecified contrasts and clustered predictive validation are described using editable mathematical equations. Rather than fabricate a dataset, the results section develops theoretical propositions, directional expectations, falsification tests and a complete reporting template. Transferable insights from cybersecurity and infrastructure-risk literature are confined to the design of decision-support and training systems; finance literature is discussed solely as an indirect implementation and resource-mobilization analogy. The report specifies conditions in which psychological safety may enable error reporting and adjustment without replacing unambiguous command authority. Its contribution is a transparent empirical architecture for future assessment of how relational climate and role clarity jointly shape adaptive emergency-response behaviour.